Population and Community Consequences of Antipredator Defenses in Ciliated Protozoans.
Ecology and Evolution of Sex Ratios in a marine copepod.

My research focusses on how adaptive changes in behaviour affect other members of a food-web through their indirect connections.
I have worked extensively on amphibian larvae and their predators in southern Michigan and on hybridogenic frogs in Europe. The work has demonstrated strong interdependencies between the availability of resources and mortality-risk for individuals and that changes in behaviour can alter the competitive regime.

The complex life histories of amphibians preclude a complete study of their population dynamics. So, I have established a model protozoan system to test the rapidly developing theories of the population and community dynamic consequences of induced anti-predator defenses.

In addition we have been investigating the evolution and ecology of biased sex ratios in the marine copepod Tigriopus californicus. This variation is heritable, and has an environmental component as well.